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Estonian Journal of Earth Sciences
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Revision of Silurian vertebrate biozones and their correlation with the conodont succession; pp. 181–204
PDF | doi: 10.3176/earth.2013.15

Authors
Tiiu Märss, Peep Männik
Abstract

The first vertebrate-based subdivisions of Silurian strata were mainly drawn on material from outcrops in Britain and drill cores from the southern Baltic. Nearly twenty years ago the first vertebrate biozonal scheme was developed on the basis of vertebrate distribution in several continuous drill core sections in the northern Baltic. This paper presents a new scheme in which many new data on vertebrate distribution from the Baltica (Baltic region, NW Russia), Avalonia (southern Britain, eastern Canada), Laurentia (northern Canada, Greenland, Scotland) and Kara (Arctic Russia) palaeocontinents have been used. All the zones have been defined, and the geographical distribution and the reference stratum and locality for each zone have been given. The Llandovery part of the succession contains the Valyalepis crista, Loganellia aldridgei and L. scotica zones; the Wenlock part is represented by the Archipelepis bifurcata/Arch. turbinata, L. grossi, Overia adraini, L. einari and Paralogania martinssoni zones. The Par. martinssoni Zone continues in the Ludlow and is followed by the Phlebolepis ornata, Phl. elegans, Andreolepis hedei, Thelodus sculptilis and T. admirabilis zones. The last zone continues in the lower Přidoli and is followed by the Nostolepis gracilis, Poracanthodes punctatus and Trimerolepis timanica zones. The L. aldridgei and Arch. bifurcata zones are new, and the Arch. turbinata and O. adraini faunas have been raised from ‘faunal succession units’ to zones. The geographically widely distributed L. grossi Zone in the upper Sheinwoodian and the Par. martinssoni Zone in the upper Homerian–lowermost Gorstian allow the integration of regional successions into one Generalized Vertebrate Zonal Scheme. Possible correlations of conodont and vertebrate biozones, and gaps in sedimentation in the northern East Baltic Silurian sequence have been discussed, the most extensive hiatus being between the Paadla and Kuressaare stages.

References

Aldridge, R. J. 1972. Llandovery conodonts from the Welsh Borderland. Bulletin of the British Museum of Natural History (Geology), 22, 127–231.

Aldridge, R. J. & Schönlaub, H. P. 1989. Conodonts. In A Global Standard for the Silurian System (Holland, C. H. & Bassett, M. G., eds), National Museum of Wales Geological Series, 9, 274–279.

Aldridge, R. J., Jeppsson, L. & Dorning, K. J. 1993. Early Silurian oceanic episodes and events. Journal of the Geological Society of London, 150, 501–513.
http://dx.doi.org/10.1144/gsjgs.150.3.0501

Aldridge, R. J., Turner, S., Jones, G. L. & Harper, D. A. T. 1996. Late Llandovery thelodonts and conodonts from the Kilbride Formation, Co. Galway, western Ireland. Geological Journal, 31, 359–367.
http://dx.doi.org/10.1002/(SICI)1099-1034(199612)31:4<359::AID-GJ713>3.3.CO;2-B

Barrick, J. E. & Klapper, G. 1976. Multielement Silurian (late Llandoverian–Wenlockian) conodonts from the Clarita Formation, Arbuckle Mountains, Oklahoma, and phylogeny of Kockelella. Geologica et Palaeontologica, 10, 59–100.

Blom, H. 1999a. Loganellia (Thelodonti, Agnatha) from the Lower Silurian of North Greenland, with special reference on their stratigraphical and palaeogeographical significance. Acta Geologica Polonica, 49, 97–104.

Blom, H. 1999b. Vertebrate remains from Upper Silurian–Lower Devonian beds of Hall Land, North Greenland. Geology of Greenland Survey Bulletin, 182, 1–80.

Blom, H. 2000. Silurian vertebrates from North Greenland. In Palaeozoic Vertebrate Biochronology and Global Marine/Non-Marine Correlation: Final Report of IGCP 328 (1991–1996) (Blieck, A. & Turner, S., eds), Courier Forschungsinstitut Senckenberg, 223, 157–160.

Calner, M. & Jeppsson, L. 2003. Carbonate platform evolution and conodont stratigraphy during the middle Silurian Mulde event, Gotland, Sweden. Geological Magazine, 140, 173–203.
http://dx.doi.org/10.1017/S0016756802007070

Corradini, C. & Serpagli, E. 1999. A Silurian biozonation from late Llandovery to end Přidoli in Sardinia (Italy). In Studies on Conodonts. Seventh European Conodont Symposium Bologna–Modena 23–25 June, 1998 (Serpagli, E. & Corradini, C., eds), Bolletino della Società Pale­onto­logica Italiana, 37, 255–273.

Cramer, B. D., Brett, C. E., Melchin, J. M., Männik, P., Kleffner, M. A., McLaughlin, P. I., Loydell, D. K., Munnecke, A., Jeppsson, L., Corradini, C., Brunton, F. R. & Saltzman, M. R. 2011. Revised correlation of Silurian Provincial Series of North America with global and regional chronostratigraphic units and δ13C chemo­stratigraphy. Lethaia, 44, 185–202.
http://dx.doi.org/10.1111/j.1502-3931.2010.00234.x

Fredholm, D. 1988. Vertebrate biostratigraphy of the Ludlovian Hemse Beds of Gotland, Sweden. Geologiska Föreningens i Stockholm Förhandlingar, 110, 237–253.
http://dx.doi.org/10.1080/11035898809452655

Fredholm, D. 1990. Agnathan vertebrates in the Lower Silurian of Gotland, Sweden. Geologiska Föreningens i Stockholm Förhandlingar, 112, 61–84.
http://dx.doi.org/10.1080/11035899009453162

Gross, W. 1967. Über Thelodontier‑Schuppen. Palaeonto­graphica, A127, 1–67.

Gross, W. 1968. Fragliche Actinopterygien-Schuppen aus dem Silur Gotlands. Lethaia, 1, 184–218.
http://dx.doi.org/10.1111/j.1502-3931.1968.tb01736.x

Helfrich, C. T. 1975. Silurian conodonts from Wills Mountain Anticline, Virginia, West Virginia and Maryland. Geological Society of America Special Paper, 161, 1–82.
http://dx.doi.org/10.1130/SPE161-p1

Jeppsson, L. 1988. Conodont biostratigraphy of the Silurian–Devonian boundary stratotype at Klonk, Czechoslovakia. Geologica et Palaeontologica, 22, 21–33.

Jeppsson, L. 1997. A new latest Telychian, Sheinwoodian and Early Homerian (early Silurian) standard conodont zonation. Transactions of the Royal Society of Edinburgh, Earth Sciences, 88, 91–114.
http://dx.doi.org/10.1017/S0263593300006854

Jeppsson, L. 2005. Conodont-based revisions of the Late Ludfordian on Gotland, Sweden. GFF, 127, 273–282.
http://dx.doi.org/10.1080/11035890501274273

Jeppsson, L. & Aldridge, R. J. 2000. Ludlow (late Silurian) oceanic episodes and events. Journal of the Geological Society, London, 157, 1137–1148.
http://dx.doi.org/10.1144/jgs.157.6.1137

Jeppsson, L., Viira, V. & Männik, P. 1994. Silurian conodont-based correlations between Gotland (Sweden) and Saaremaa (Estonia). Geological Magazine, 131, 201–218.
http://dx.doi.org/10.1017/S0016756800010736

Jeppsson, L., Eriksson, M. E. & Calner, M. 2006. A latest Llandovery to latest Ludlow high-resolution biostrati­graphy based on the Silurian of Gotland – a summary. GFF, 128, 109–114.
http://dx.doi.org/10.1080/11035890601282109

Kaljo, D., Kiipli, T. & Martma, T. 1997. Carbon isotope event markers through the Wenlock–Přidoli sequence at Ohesaare (Estonia) and Priekule (Latvia). Palaeogeography, Palaeo­climatology, Palaeoecology, 132, 211–223.
http://dx.doi.org/10.1016/S0031-0182(97)00065-5

Karatajūtė-Talimaa, V. 1978. Telodonty silura i devona SSSR i Spitsbergena [Thelodonts from the Silurian and Devonian of the USSR and Spitsbergen]. Mokslas, Vilnius, 334 pp. [in Russian].

Karatajūtė‑Talimaa, V. & Brazauskas, A. 1995. Distribution of vertebrates in the Silurian of Lithuania. Geologija (Vilnius), 17, 106–114.

Karatajūtė-Talimaa, V. & Märss, T. 2002. Upper Silurian thelodonts of Severnaya Zemlya Archipelago (Russia). Geodiversitas, 24, 405–443.

Karatajūtė-Talimaa, V. N., Valiukevičius, J. J. & Brazauskas, A. Z. 1987. Rasprostranenie konodontov i pozvonochnykh v silurijskikh otlozheniyakh Litvy [The distribution of conodonts and vertebrates in the Silurian deposits of Lithuania]. Geologija (Vilnius), 8, 59–71 [in Russian].

Loydell, D. K., Männik, P. & Nestor, V. 2003. Integrated biostratigraphy of the lower Silurian of the Aizpute-41 core, Latvia. Geological Magazine, 140, 205–229.
http://dx.doi.org/10.1017/S0016756802007264

Loydell, D. K., Nestor, V. & Männik, P. 2010. Integrated bio­stratigraphy of the lower Silurian of the Kolka-54 core, Latvia. Geological Magazine, 147, 253–280.
http://dx.doi.org/10.1017/S0016756809990574

Männik, P. 1999. Ordovician and Silurian conodonts. In Silurian and Devonian Stratigraphy of the Severnaya Zemlya Archipelago (Matukhin, R. G. & Menner, V. Vl., eds), pp. 110–120. SNIIGiMS, Novosibirsk.

Männik, P. 2001. Evolution of early Silurian conodont faunas, and high-resolution stratigraphy. In Evolution of Life on the Earth: Proceedings of the II International Symposium ‘Evolution of Life on the Earth’, November 12–15, 2001, Tomsk (Podobina, V. M., Rodygin, S. A., Savina, N. I. & Tat´yanin, G. M., eds), pp. 202–205. Tomsk State University, Tomsk.

Männik, P. 2002. Conodonts in the Silurian of Severnaya Zemlya and Sedov archipelagos (Russia), with special reference to the genus Ozarkodina Branson & Mehl, 1993. Geodiversitas, 24, 77–97.

Männik, P. 2007a. An updated Telychian (Late Llandovery, Silurian) conodont zonation based on Baltic faunas. Lethaia, 40, 45–60.
http://dx.doi.org/10.1111/j.1502-3931.2006.00005.x

Männik, P. 2007b. Some comments on the Telychian–early Sheinwoodian conodont faunas, events and stratigraphy. Acta Palaeontologica Sinica, 46 (Suppl.), 305–310.

Männik, P. & Małkowski, K. 1998. Silurian conodonts from the Gołdap core, Poland. In Proceedings of the Sixth European Conodont Symposium (ECOS VI) (Szaniawski, H., ed.), Palaeontologica Polonica, 58, 139–149.

Männik, P. & Martma, T. 2000. Llandovery–Wenlock boundary in the Subpolar Urals. In Pan-Arctic Palaeozoic Tectonics, Evolution of Basins and Faunas (Antoshkina, A., Malysheva, E. & Wilson, M. V. H., eds), Ichthyolith Issues Special Publication, 6, 64–67.

Männik, P., Antoshkina, A. I. & Beznosova, T. M. 2000. The Llandovery–Wenlock boundary in the Russian Arctic. Proceedings of the Estonian Academy of Sciences, Geology, 49, 104–111.

Männik, P., Bogolepova, O. K., Põldvere, A. & Gubanov, A. P. 2009. New data on Ordovician–Silurian conodonts and stratigraphy from the Severnaya Zemlya archipelago, Russian Arctic. Geological Magazine, 146, 497–519.
http://dx.doi.org/10.1017/S0016756809006372

Märss, T. 1982a. Vertebrate zones in the East Baltic Silurian. In Ecostratigraphy of the East Baltic Silurian (Kaljo, D. & Klaamann, E., eds), pp. 97–106. Valgus, Tallinn.

Märss, T. 1982b. Biozones of vertebrates (Silurian of North Baltic). In Soobshchestva i biozony v silure Pribaltiki [Communities and Biozones in the Baltic Silurian] (Kaljo, D. & Klaamann, E., eds), pp. 97–116. Valgus, Tallinn [in Russian].

Märss, T. 1986. Silurian vertebrates of Estonia and West Latvia. Fossilia Baltica, 1, 1–104 [in Russian, with English summary].

Märss, T. 1989. Vertebrates. In A Global Standard for the Silurian System (Holland, C. H. & Bassett, M. G., eds), National Museum of Wales, Geological Series, 9, 284–289.

Märss, T. 1990. Vertebrates. In Field Meeting, Estonia 1990: An Excursion Guidebook (Kaljo, D. & Nestor, H., eds), pp. 93–96. Estonian Academy of Sciences, Tallinn.

Märss, T. 1992. Vertebrate history in the Late Silurian. Proceedings of the Estonian Academy of Sciences, Geology, 41, 205–214.

Märss, T. 1996. Loganellia (Thelodonti, Agnatha) from the Jaagarahu Stage, Wenlock, Estonia. Proceedings of the Estonian Academy of Sciences, Geology, 45, 189–202.

Märss, T. 1997. Vertebrates of the Přidoli and Silurian–Devonian boundary beds in Europe. Modern Geology, 21, 17–42.

Märss, T. 2000. Silurian vertebrate studies during 1990–1996. In Palaeozoic Vertebrate Biochronology and Global Marine/Non-Marine Correlation: Final Report of IGCP 328 (1991–1996) (Blieck, A. & Turner, S., eds), Courier Forschungsinstitut Senckenberg, 223, 81–90.

Märss, T. 2001. Andreolepis (Actinopterygii) in the Upper Silurian of northern Eurasia. Proceedings of the Estonian Academy of Sciences, Geology, 50, 190–206.

Märss, T. 2003. Paralogania from the Rootsiküla (Wenlock) and Paadla (Ludlow) stages of Estonia. Proceedings of the Estonian Academy of Sciences, Geology, 52, 98–112.

Märss, T. 2006. Thelodonts (Agnatha) from the basal beds of the Kuressaare Stage, Ludlow, Upper Silurian of Estonia. Proceedings of the Estonian Academy of Sciences, Geology, 55, 43–66.

Märss, T. & Karatajūtė-Talimaa, V. 2002. Ordovician and Lower Silurian thelodonts from Severnaya Zemlya Archipelago (Russia). Geodiversitas, 24, 381–404.

Märss, T. & Miller, C. G. 2004. Thelodonts and distribution of associated conodonts from the Wenlock–Lowermost Lochkovian of the Welsh Borderland. Palaeontology, 47, 1211–1265.
http://dx.doi.org/10.1111/j.0031-0239.2004.00409.x

Märss, T. & Ritchie, A. 1998. Articulated thelodonts (Agnatha) of Scotland. Transactions of the Royal Society of Edinburgh, Earth Sciences, 88, 143–195.
http://dx.doi.org/10.1017/S026359330000691X

Märss, T., Fredholm, D., Talimaa, V., Turner, S., Jeppsson, L. & Nowlan, G. 1995. Silurian Vertebrate Biozonal Scheme. In Premiers Vertébrés et Vertébrés Inferieurs (Lelievre, H., Wenz, S., Blieck, A. & Cloutier, R., eds), Geobios, M.S., 19, 368–372.

Märss, T., Fredholm, D., Talimaa, V., Turner, S., Jeppsson, L. & Nowlan, G. 1996. Towards the Silurian Vertebrate Biozonal Standard. In The James Hall Symposium: Second International Symposium on the Silurian System; Program and Abstracts (Brett, C. E., ed.), pp. 73–74. University of Rochester, N.Y., August 4–9, 1996.

Märss, T., Caldwell, M., Gagnier, P.-Y., Goujet, D., Männik, P., Martma, T. & Wilson, M. 1998a. Distribution of Silurian and Lower Devonian vertebrate microremains and conodonts in the Baillie-Hamilton and Cornwallis Island sections, Canadian Arctic. Proceedings of the Estonian Academy of Sciences, Geology, 47, 51–76.

Märss, T., Soehn, K. L. & Wilson, M. V. H. 1998b. Micro­vertebrate-based correlations of the Llandovery–Wenlock boundary in some sections of the Selwyn and Franklinian sedimentary basins, northern Canada. In Circum-Arctic Palaeozoic Faunas and Facies (Ginter, M. & Wilson, M. V. H., eds), Ichthyolith Issues Special Publications, 4, 37–39.

Märss, T., Wilson, M. V. H. & Thorsteinsson, R. 2002. New thelodont (Agnatha) and possible chondrichthyan (Gnathostomata) taxa established in the Silurian and Lower Devonian of Arctic Canada. Proceedings of the Estonian Academy of Sciences, Geology, 51, 88–120.

Märss, T., Wilson, M. V. H. & Thorsteinsson, R. 2006. Silurian and Lower Devonian thelodonts and putative chondrichthyans from the Canadian Arctic Archipelago. Special Papers in Palaeontology, 75, 1–140.

Märss, T., Turner, S. & Karatajūtė-Talimaa, V. 2007. ‘Agnatha’ II. Thelodonti. In Handbook of Paleoichthyology, Vol. 1B (Schultze, H.-P., ed.). Verlag Dr. Friedrich Pfeil, München, 143 pp.

Matukhin, R. G., Menner, V. V. & Kuršs, V. M. 1999. Structure of the sections. In Biostratigrafiya silura i devona arkhipelaga Severnaya Zemlya [Biostratigraphy of the Silurian and Devonian of the Severnaya Zemlya Archipelago] (Matukhin, R. G. & Menner, V. V., eds), pp. 7–82. SNIIGiMS, Novosibirsk [in Russian].

Miller, G. & Märss, T. 1999. A conodont, thelodont and acanthodian fauna from the Lower Přidoli (Silurian) of the Much Wenlock Area, Shropshire. Palaeontology, 42, 691–784.
http://dx.doi.org/10.1111/1475-4983.00093

Modzalevskaya, T. & Märss, T. 1991. On the age of the lower boundary of the Greben Regional Stage of the Urals. Proceedings of the Estonian Academy of Sciences, Geology, 40, 100–103 [in Russian, with English summary].

Nestor, H. 1997. Silurian. In Geology and Mineral Resources of Estonia (Raukas, A. & Teedumäe, A., eds), pp. 89–106. Estonian Academy Publishers, Tallinn.

Nestor, H., Einasto, R., Nestor, V., Märss, T. & Viira, V. 2001. Description of the type section, cyclicity, and correlation of the Riksu Formation (Wenlock, Estonia). Proceedings of the Estonian Academy of Sciences, Geology, 50, 149–173.

Nestor, H., Einasto, R., Männik, P. & Nestor, V. 2003. Correlation of some lower–middle Llandovery reference sections in central and southern Estonia and sedimen­tation cycles of lime muds. Proceedings of the Estonian Academy of Sciences, Geology, 52, 3–27.

Nowlan, G. S. (ed.). 1995. Left hand column for correlation charts. Silurian Times. A Newsletter of the Silurian Subcommission. 3, 7–11. Geological Survey of Canada, Calgary.

Over, D. J. & Chatterton, B. D. E. 1987. Silurian conodonts from the southern Mackenzie Mountains, Northwest Territories, Canada. Geologica et Palaeontologica, 21, 1–49.

Plax, D. & Märss, T. 2011. Thelodonts (Agnatha) from the Lower Silurian (Wenlock) deposits of the northwest of Belarus. Litasfera [Lithosphere], 34, 69–81.

Ritchie, A. 1985. Ainiktozoon loganense Scourfield, a proto­chordate from the Silurian of Scotland. Alcheringa, 9, 117–142.
http://dx.doi.org/10.1080/03115518508618961

Shuiskij, V. P. 1981. Facial-lithological peculiarities of the Upper Silurian and Lower Devonian reefs on the western slope of the Central Urals. In Biostratigrafiya i fauna srednego paleozoya Urala [Biostratigraphy and Fauna of the Middle Palaeozoic of the Urals] (Sapel¢nikov, V. P. & Chuvashov, B. I., eds), pp. 96–110. IGiG UNTs AN SSSR, Sverdlovsk [in Russian].

Shurygina, M. V., Breivel¢, M. G., Breivel¢, I. A., Zenkova, G. G., Militsina, V. S. & Yanet, F. E. 1981. Přidoli on the North and Central Urals. In Biostratigrafiya i fauna srednego paleozoya Urala [Biostratigraphy and Fauna of the Middle Palaeozoic of the Urals] (Sapel¢nikov, V. P. & Chuvashov, B. I., eds), pp. 55–74. IGiG UNTs AN SSSR, Sverdlovsk [in Russian].

Soehn, K. L., Hanke, G. F., Märss, T. & Wilson, M. V. H. 2000. Preliminary vertebrate biostratigraphy of the Avalanche Lake sections (Wenlock, Silurian), southern Mackenzie Mountains, N.W.T., and review of north­western Canadian vertebrate localities of Silurian Age. In Palaeozoic Vertebrate Biochronology and Global Marine/Non-Marine Correlation: Final Report of IGCP 328 (1991–1996) (Blieck, A. & Turner, S., eds), Courier Forschungsinstitut Senckenberg, 223, 129–156.

Soehn, K. L., Märss, T., Caldwell, M. W. & Wilson, M. V. H. 2001. New and biostratigraphically useful thelodonts from the Silurian of the Mackenzie Mountains, Northwest Territories, Canada. Journal of Vertebrate Paleontology, 21, 651–659.
http://dx.doi.org/10.1671/0272-4634(2001)021[0651:NABUTF]2.0.CO;2

Talimaa, V. 2000. Significance of thelodonts (Agnatha) in correlation of the Upper Ordovician to Lower Devonian of the northern part of Eurasia. In Palaeozoic Vertebrate Biochronology and Global Marine/Non-Marine Correlation: Final Report of IGCP 328 (1991–1996) (Blieck, A. & Turner, S., eds), Courier Forschungsinstitut Senckenberg, 223, 69–80.

Turner, S. 1973. Siluro-Devonian thelodonts from the Welsh Borderland. Journal of the Geological Society, 129, 557–584.
http://dx.doi.org/10.1144/gsjgs.129.6.0557

Turner, S. 1984. Studies on Palaeozoic Thelodonti (Craniata: Agnatha). Unpublished PhD thesis (2 vols), 272 pp., University of Newcastle-upon-Tyne.

Turner, S. 2000. New Llandovery to early Přidoli micro­vertebrates including Lower Silurian zone fossil, Loganellia avonia nov. sp., from Britain. In Palaeozoic Vertebrate Biochronology and Global Marine/Non-Marine Correlation: Final Report of IGCP 328 (1991–1996) (Blieck, A. & Turner, S., eds), Courier Forschungsinstitut Senckenberg, 223, 91–127.

Turner, S. & Nowlan, G. S. 1995. Early Silurian micro­vertebrates of eastern Canada. Bulletin du Museum national d’Histoire naturelle, 17, 513–529.

Turner, P. & Turner, S. 1974. Thelodonts from the Upper Silurian of Ringerike, Norway. Norsk Geologisk Tiddskrift, 54, 182–192.

Uyeno, T. T. 1990. Biostratigraphy and conodont faunas of Upper Ordovician through Middle Devonian rocks, eastern Arctic Archipelago. Bulletin of the Geological Survey of Canada, 401, 1–211.
http://dx.doi.org/10.4095/129059

Valiukevičius, J. 2003. New Silurian nostolepids (Acanthodii, Pisces) of Lithuania. Geologija (Vilnius), 42, 51–68.

Valiukevičius, J. 2004a. Silurian acanthodian succession of the Lužni-4 borehole (Latvia). Acta Universitatis Latviensis, 679, 120–147.

Valiukevičius, J. 2004b. New Wenlock–Přidoli (Silurian) acanthodian fishes from Lithuania. Acta Palaeontologica Polonica, 49, 147–160.

Valiukevičius, J. 2005. Silurian acanthodian biostratigraphy of Lithuania. Geodiversitas, 27, 349–380.

Valiukevičius, J. 2006. Event pattern in the development of Siluro-Devonian acanthodians of Lithuania. Neues Jahrbuch der Geologie und Paläontologie, 6, 321–343.

Valiukevičius, J., Gladkovskij, V. T., Karatajūtė-Talimaa, V. N., Kuršs, V., Mel¢nikov, S. V. & Menner, V. V. 1983. Stratigrafiya silura i nizhnego devona Severnogo Timana [Stratigraphy of the Silurian and Lower Devonian of Northern Timan]. Seriya Geologicheskaya, 10, 53–64 [in Russian].

Vergoossen, J. M. J. 1999. Siluro-Devonian microfossils of Acanthodii and Chondrichthyes (Pisces) from the Welsh Borderland/south Wales. Modern Geology, 24, 23–90.

Viira, V. 1982a. Late Silurian shallow and deep water cono­donts of the East Baltic. In Ecostratigraphy of the East Baltic Silurian (Kaljo, D. & Klaamann, E., eds), pp. 79–87. Valgus, Tallinn.

Viira, V. 1982b. Shallow-water conodont Ctenognathodus murchisoni (Late Wenlock, Estonia). In Communities and Biozones in the Baltic Silurian (Kaljo, D. & Klaamann, E., eds), pp. 63–83. Valgus, Tallinn.

Viira, V. 1999. Late Silurian conodont biostratigraphy in the northern East Baltic. In Studies on Conodonts. Seventh European Conodont Symposium Bologna–Modena 23–25 June, 1998 (Serpagli, E. & Corradini, C., eds), Bolletino della Società Paleontologica Italiana, 37, 299–310.

Viira, V. 2000. Latest Silurian (Ohesaare Stage) conodonts and the detorta Zone in the northern East Baltic. Proceedings of the Estonian Academy of Sciences, Geology, 49, 44–62.

Viira, V. & Aldridge, R. J. 1998. Upper Wenlock to lower Přidoli (Silurian) conodont biostratigraphy of Saaremaa, Estonia, and a correlation with Britain. Journal of Micro­palaeontology, 17, 33–50.
http://dx.doi.org/10.1144/jm.17.1.33

Viira, V. & Männik, P. 1997. Conodonts. In Geology and Mineral Resources of Estonia (Raukas, A. & Teedumäe, A., eds), pp. 241–244. Estonian Academy Publishers, Tallinn.

Walliser, O. H. 1964. Conodonten des Silurs. Abhandlungen des Hessischen Landesamtes für Bodenforschung, 41, 1–106.

Žigaitė, Ž. & Blieck, A. 2006. Palaeobiogeographical significance of Early Silurian thelodonts from central Asia and southern Siberia. GFF, 128, 203–206.
http://dx.doi.org/10.1080/11035890601282203

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