eesti teaduste
akadeemia kirjastus
SINCE 1952
Earth Science cover
Estonian Journal of Earth Sciences
ISSN 1736-7557 (Electronic)
ISSN 1736-4728 (Print)
Impact Factor (2021): 0.811
Neotypes for some upper Silurian acanthodian taxa from the Baltic Sea Region and the Welsh Borderland; pp. 17–24
PDF | 10.3176/earth.2022.02

Carole J. Burrow, Tiiu Märss

The type material of acanthodian taxa from the upper Silurian of Estonia, erected by C. Pander and V. Rohon in the nineteenth century, has long been lost, and in most cases no neotypes have been erected to date. We nominate scales of Nostolepis striata Pander and Gomphonchus sandelensis (Pander) from the type locality at the Ohesaare Cliff, Saaremaa Island, Estonia, as neotypes for those two species. We also nominate a replacement neotype scale for Radioporacanthodes biblicus (Lehman) from the vicinity of the type locality at Ramsåsa, southern Sweden, to replace the invalid neotype from a borehole on Saaremaa Island. The validity and distribution of Gomphonchus volborthi (Rohon) is considered, and a neotype and type stratum (the Ludlow Bone Bed, Shropshire) are designated. Specimen repository details, where not provided in the original descriptions, are listed for other acanthodian scale-based taxa from the Baltic Sea Region.


Bremer, O., Niedźwiedzki, G., Blom, H., Dec, M. & Kozłowski, W. 2018. Vertebrate microremains from the upper Silurian Winnica Formation of the Holy Cross Mountains, Poland. Geological Magazine155, 1523–1541.

Bremer, O., Turner, S., Märss, T. & Blom, H. 2019a. Silurian vertebrate remains from the Oslo Region, Norway, and their implications for regional biostratigraphy. Norwegian Journal of Geology99, 129–150 + 5 appendices.

Bremer, O., Jarochowska, E. & Märss, T. 2019b. Vertebrate remains and conodonts in the upper Silurian Hamra and Sundre formations of Gotland, Sweden. GFF142, 52–80.

Brotzen, F. 1934. Erster Nachweis von Unterdevon im Ostseegebiete durch Konglomeratgeschiebe mit Fischresten. II Teil (Paläontologie). Zeitschrift Geschiebeforschung10, 1–65.

Burrow, C. J. 2021. Acanthodii, Stem Chondrichthyes. In Handbook of Paleoichthyology, Volume 5 (Schultze, H.-P., ed.). Verlag Dr Friedrich Pfeil, Munich, 135 pp. 

Burrow, C. J. & Turner, S. 2018. Stem chondrichthyan microfossils from the Lower Old Red Sandstone of the Welsh Borderland. Acta Geologica Polonica68, 321–334.

Catlos, E. J., Mark, D. F., Suarez, S., Brookfield, M. E., Miller, C. G., Schmitt, A. K., Gallagher, V. & Kelly, A. 2020. Late Silurian zircon U–Pb ages from the Ludlow and Downton bone beds, Welsh Basin, UK. Journal of the Geological Society178, jgs2020-107.

Coates, M. I., Finarelli, J. A., Sansom, I. J., Andreev, P. S., Criswell, K. E., Tietjen, K., Rivers, M. L. & La Riviere, P. J. 2018. An early chondrichthyan and the evolutionary assembly of a shark body plan. Proceedings of the Royal Society B: Biological Sciences285, 20172418.

Denison, R. H. 1979. Acanthodii. In Handbook of Paleoichthyology, Volume 5 (Schultze, H.-P., ed.). Gustav Fischer Verlag, Stuttgart and New York, 62 pp.

Fredholm, D. 1988. Vertebrates in the Ludlovian Hemse Beds of Gotland, Sweden. Geologiska Foreningens i Stockholm Forhandlingar110, 157–179.

Gross, W. 1947. Die Agnathen und Acanthodier des ober­silurischen Beyrichienkalks. Palaeontographica Abt. A96, 91–161.

Gross, W. 1971. Downtonische und Dittonische Acanthodier-Reste des Ostseegebietes. Palaeontographica Abt. A136, 1–82.

Hoppe, K. H. 1931. Die Coelepiden und Acanthodier des Obersilurs der Insel Oesel. Ihre Paläobiologie und Paläontologie. Palaeontographica Abt. A76, 38–94.

ICZN. 1999. International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, UK, 306 pp.

Lehman, J.-P. 1937. Les poissons du Downtonien de la Scanie (Suède). Mémoires du Faculté Sciences, Université de Paris664, 1–98. 

Mark-Kurik, E., Märss, T. & Kuršs, V. 1989. Excursion Guidebook. The Silurian of Saaremaa and the Devonian of South Estonia and North Latvia. 2nd International Colloquium on the Middle Palaeozoic Fishes. Tallinn, September 1989. Estonian Academy of Sciences Editorial and Publishing Council, Tallinn, 43 pp.

Märss, T. 1986. Silurian vertebrates of Estonia and West Latvia. Fossilia Baltica1, 1–104 [in Russian, with English summary].

Pander, C. H. 1856. Monographie der fossilen Fische des silurischen Systems der Russisch-Baltischen Gouvernements. Obersilurische Fische. Buchdruckerei der Kaiserlichen Akademie der Wissenschaften, St. Petersburg, 91 pp.

Plax, D. P. 2019. On the redeposited Silurian ichthyofauna remains in the Quaternary deposits of Belarus. Natural Resources, 2/2019, 34–45.

Reuss, A. E. 1846. Die Versteinerungen der böhmischen Kreideformation. Zweite Abtheilung. E. Schweizerbart’sche Verlagsbuchhandlung, Stuttgart, iv + 148 pp.

Rohon, J. von. 1893. Die obersilurischen Fische von Oesel. Teil II. Selachii, Dipnoi, Ganoidei, Pteraspidae und Cephalaspidae. Memoires de l’Académie Impériale des Sciences de St. Petersbourg, VIIe série, 41, 1–124.

Valiukevičius, J. J. 1998. Acanthodians and zonal stratigraphy of Lower and Middle Devonian in East Baltic and Byelorussia. Palaeontographica A248, 1–53.

Valiukevičius, J. J. 2003a. New Silurian nostolepids (Acanthodii, Pisces) of Lithuania. Geologija (Vilnius)42, 51–68.

Valiukevičius, J. J. 2003b. New Late Silurian to Middle Devonian acanthodians of the Timan-Pechora region. Acta Geologica Polonica53, 209–245.

Valiukevičius, J. 2004. Silurian succession of the Luzni-4 borehole (Latvia). Acta Universitatis Latviensis679, 120–147.

Valiukevičius, J. & Burrow, C. J. 2005. Diversity of tissues in acanthodians with “Nostolepis”-type histological structure. Acta Palaeontologica Polonica50, 635–649.

Vergoossen, J. M. J. 1997. Revision of poracanthodid acanthodians. Ichthyolith Issues Special Publication3, 44–46.

Vergoossen, J. M. J. 1999a. Siluro-Devonian microfossils of Acanthodii and Chondrichthyes (Pisces) from the Welsh Borderland/south Wales. Modern Geology24, 23–90.

Vergoossen, J. M. J. 1999b. Late Silurian fish microfossils from Helvetesgraven, Skane (southern Sweden) (I). Geologie en Mijnbouw78, 267–280.

Vergoossen, J. M. J. 1999c. Late Silurian fish microfossils from an East Baltic-derived erratic from Oosterhaule, with a description of new acanthodian taxa. Geologie en Mijnbouw78, 231–251.

Vergoossen, J. M. J. 2000. Acanthodian and chondrichthyan microremains in the Siluro-Devonian of the Welsh Borderland, Great Britain, and their biostratigraphical potential. Courier Forschungsinstitut Senckenberg223, 175–199.

Vergoossen, J. M. J. 2002. Late Silurian fish microfossils from Ramsåsa, Locality H, Scania, south Sweden, with some remarks on the body zonation scheme used in thelodont studies. Scripta Geologica123, 41–69.

Vergoossen, J. M. J. 2003. First record of fish microfossils from Ramsåsa, site C, Skåne, southern Sweden. Scripta Geologica126, 1–78.

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