ESTONIAN ACADEMY
PUBLISHERS
eesti teaduste
akadeemia kirjastus
PUBLISHED
SINCE 1952
 
Earth Science cover
Estonian Journal of Earth Sciences
ISSN 1736-7557 (Electronic)
ISSN 1736-4728 (Print)
Impact Factor (2022): 1.1
Research article
Taxonomy of the Sandbian (Upper Ordovician) brachiopod Dalmanella kegelensis Alichova, 1953 and the new genus Alichovella; pp. 45–56
PDF | https://doi.org/10.3176/earth.2024.06

Author
Linda Hints
Abstract

The brachiopod species Dalmanella kegelensis Alichova, 1953 is reported in the geological literature of the Baltic region as a species characteristic of the Keila Regional Stage at the Sandbian–Katian transition, and a nominal taxon of a regional biozone. The spatial distribution of D. kegelensis forms a belt around the deeper parts of the palaeobasin. The easternmost occurrences of D. kegelensis have been recorded at the westernmost periphery of the Moscow Basin. The selection of the holotype of the species D. kegelensis from a locality in western Estonia caused some confusion in understanding Alichova’s species, since the description of the species is mainly based on brachiopods collected from northwestern Russia. The earlier attempt to revise the taxonomy of D. kegelensis group brachiopods was unsustainable, while assigning these brachiopods to the genus Horderleyella is no longer feasible. However, two subspecies were identified, which are discussed here as two separate species, kegelensis and oanduensis, belonging to two different genera: one species belongs to the new genus Alichovella in the family Draboviidae, and the second one, following Alichova’s classification, to the genus Dalmanella in the family Dalmanellidae. These species possibly represent somewhat different ages prior to the beginning of the Guttenberg carbon isotope excursion and supposedly differ in their positions on the onshore-offshore transect.

References

Ainsaar, L. and Meidla, T. 2001. Facies and stratigraphy of the middle Caradoc mixed siliciclastic-carbonate sediments in eastern Baltoscandia. Proceedings of the Estonian Academy of Sciences, Geology50, 5–23.
https://doi.org/10.3176/geol.2001.1.02

Ainsaar, L., Meidla, T. and Martma, T. 1999. Evidence for a widespread carbon isotopic event associated with late Middle Ordovician sedimentological and faunal changes in Estonia. Geological Magazine, 136(1), 49–62.
https://doi.org/10.1017/S001675689900223X

Ainsaar, L., Kaljo, D., Martma, T., Meidla, T., Männik, P., Nõlvak, J., and Tinn, O. 2010. Middle and Upper Ordovician carbon isotope chemostratigraphy in Baltoscandia: a correlation standard and clues to environmental history. Palaeogeography, Palaeoclimatology, Palaeoecology, 294(3–4), 189–201. 
https://doi.org/10.1016/j.palaeo.2010.01.003  

Alichova, T. N. 1951. Брахиоподы средней и верхней части нижнего силура Ленинградской области и их cтратиграфи­ческое значение (Brachiopods of the Middle and Upper Parts of the Lower Silurian in Leningrad District and Their Stratigraphic Meaning). Госгеолиздат, Москва.

Alichova, T. N. 1953. Руководящая фауна брахиопод ордовикских отложений Северо-Западной части Русской платформы (Index Brachiopod Fauna from the Ordovician Deposits of the North-Western Part of the Russian Platform). ВСЕГЕИ (VSEGEI), State Publishing House of Geological Literature, St. Petersburg.

Alichova, T. N. 1960. Стратиграфия ордовикских отложений Русской платформы (Stratigraphy of the Ordovician Deposits of the Russian Platform).ВСЕГЕИ (VSEGEI), State Scientific and Technical Publishing House of Literature on Geology, Geodesy and Subsoil Protection, Moscow.

Alichova, T. N. 1969. Стратиграфия и брахиоподы среднеордо­викских отложений Московской синеклизы (Stratigraphy and Brachiopods of the Middle Ordovician Deposits in the Moscow Syneclise). Nedra, Moscow.

Alichova, T. N., Balashova, E. A. and Balashov, Z. G. 1954. Полевой атлас характерных комплексов фауны отложений ордовика и готландия южной части Литовской ССР (Field Atlas of Characteristic Complexes of Ordovician and Gotlandian Faunas of the Southern Lithuanian SSR). Nedra, Moscow.

Bauert, H., Hints, O., Bauert, C., Nõlvak, J., Ainsaar, L. and Martma, T. 2017. The Guttenberg carbon isotope excursion (GICE; Ordovician) in Estonia. In Proceedings of the 10th Baltic Stratigraphic Conference, Chęciny, 12–14 September 2017 (Źylińska, A., ed.). University of Warsaw, Warsaw, 13–14.

Bergström, S. M., Huff, W. D., Kolata, D. R. and Bauert, H. 1995. Nomenclature, stratigraphy, chemical fingerprinting and areal distribution of some Middle Ordovician K-bentonites in Balto­scandia. GFF117, 1–13.
https://doi.org/10.1080/11035899509546191

Bergström, S. M., Schmitz, B., Young, S. A. and Bruton, D. L. 2011. Lower Katian (Upper Ordovician) δ13C chemostratigraphy, global correlation and sea-level changes in Baltoscandia. GFF133, 31–47.
https://doi.org/10.1080/11035897.2011.557162

Candela, Y. 2003. Late Ordovician Brachiopods from the Bardahes­siagh Formation of Pomeroy, Ireland. Monographs of the Palae­onto­­graphical Society, London.

Cocks, L. R. M. 2008. A Revised Review of British Lower Palaeo­zoic Brachiopods. Monographs of the Palaeontological Society, London.

Cooper, G. A. 1930. New species from the Upper Ordovician of Perce. American Journal of Science, 20(119), 265–292.

Cooper, G. A. 1956. Chazyan and related brachiopods. Smithsonian Miscellaneous Collections, 127(1), 1–1245.

Dmitrovskaya, J. E. 1991. Нижний палеозой Московской синеклизы (The Lower Paleozoic of the Moscow Syneclise). PhD thesis, Moscow.

Gagel, C. 1890. Die Brachiopoden der cambrischen und silurischen Geschiebe im Diluvium der Provinzen Ost- und WestpreussenKoch, Königsberg.

Harper, D. A. T. 1984. Brachiopods from the Upper Ardmillan Succession (Ordovician) of the Girvan District, Scotland. Part I. Monographs of the Palaeontographical Society, London.

Harris, M. T., Sheehan, P. M., Ainsaar, L., Hints, L., Männik, P., Nõlvak, J. and Rubel, M. 2004. Upper Ordovician sequences of western Estonia. Palaeogeography, Palaeoclimatology, Palaeoecology, 210(2–4), 135–148.
https://doi.org/10.1016/j.palaeo.2004.02.045

Havlíček, V. 1950. Ramenonožci českého ordoviku (The Ordovician brachiopoda from Bohemia). Rozpravy Ústředního ústavu geologického, 13, 1–72.

Hints, L. 1975. Брахиоподы Enteletacea ордовика Прибалтики (Ordovician Brachiopods Enteletacea of the East Baltic Area). Academy of Sciences of the Estonian SSR, Tallinn.

Hints, L. 2012. New Hirnantian orthide brachiopods from the type section of the Porkuni Stage (Porkuni quarry, northeastern Estonia). Estonian Journal of Earth Sciences, 61(4), 227–241.
https://doi.org/10.3176/earth.2012.4.04

Hints, L. and Nõlvak, J. 2023. Latest Sandbian brachiopods and chitinozoan biostratigraphy in North Estonia. Estonian Journal of Earth Sciences, 72(1), 42–45.
https://doi.org/10.3176/earth.2023.50

Hints, L., Paškevičius, J., Martma, T., Männik, P. and Nõlvak, J. 2016. Upper Sandbian–lower Katian bio- and chemostratigraphy in the Pajevonys-13 core section, Lithuania. Estonian Journal of Earth Sciences, 65(2), 85–97.
https://doi.org/10.3176/earth.2016.08

Hurst, J. M. 1979. The stratigraphy and brachiopods of the upper part of the type Caradoc of south Salop. Bulletin of the British Museum (Natural History), Geology, 32(10), 183–304.

ICZN (International Commission on Zoological Nomenclature) 2000. International Code of Zoological Nomenclature. 4th ed. The International Trust for Zoological Nomenclature, London.

Jin, J. 2012. Cincinnetina, a new Late Ordovician dalmanellid brachiopod from the Cincinnati type area, USA: implications for the evolution and palaeogeography of the epicontinental fauna of Laurenia. Palaeontology, 55(1), 205–228.
https://doi.org/10.1111/j.1475-4983.2011.01113.x

Jin, J. and Bergström, J. 2010. True Dalmanella and taxonomic implications for some Late Ordovician dalmanellid brachiopods from North America. GFF,132(1), 13–24.
https://doi.org/10.1080/11035891003597083

Kröger, B., Hints, L. and Lehnert, O. 2014. Age, facies, and geometry of the Sandbian/Katian (Upper Ordovician) pelma­tozoan-bryozoan-receptaculitid reefs of the Vasalemma Formation, northern Estonia. Facies60, 963–986.
https://doi.org/10.1007/s10347-014-0410-8

Ludkevich, E. M. 1939. Иевский ярус силурийского плато Прибалтики (The Jewe Stage of the East Baltic Silurian Plateau). Bulletin of the Imperial Society of Naturalists of Moscow, Geological Series, 17(4–5), 135–142.

Ludvigson, G. A., Jacobson, S. R., Witzke, B. J. and González, L. A. 1996. Carbonate component chemostratigraphy and depositional history of the Ordovician Decorach Formation, Upper Mississippi Valley. Geological Society of America Special Paper, 306, 67–86.
https://doi.org/10.1130/0-8137-2306-X.67

Männil, R. M. 1958. Основные черты стратиграфии кейлаского горизонта (DII, oрдовик) в Эстонии (Grundzüge der Strati­graphie der Keila-Stufe (Ordovizium, Estland)). Eesti NSV Teaduste Akadeemia toimetised, Tehniliste ja füüsikalis-matemaatiliste teaduste seeria, 7(3), 235–246.
https://doi.org/10.3176/tech.phys.math.1958.3.08

Männil, R. and Meidla, T. 1994. The Ordovician System of the East European Platform (Estonia, Latvia, Lithuania, Byelorussia, parts of Russia, the Ukraine and Moldova). In The Ordovician System of the East European Platform and Tuva (Southeastern Russia): Correlation charts and explanatory notes (Webby, B. D., Ross, R. J., Jr. and Zhen, Y. Y., eds). International Union of Geo­logical Sciences Publication, 28, 1–52.

Meidla, T., Ainsaar, L., Hints, L., Hints, O., Martma, T. and Nõlvak, J. 1999. The mid-Caradocian biotic and isotopic event in the Ordovician of the East Baltic. Acta Universtatis Carolinae, Geologica, 43(1/2), 503–506.

Mitchell, W. I. 1977. The Ordovician Brachiopoda from Pomeroy, Co. Tyrone. Monographs of the Palaeontographical Society, London.
https://doi.org/10.1080/25761900.2022.12131731

Moore, R. C. 1952. Brachiopoda. In Invertebrate Fossils (Moore, R. C., Lalicker, C. G. and Fischer, A. G., eds). McGraw-Hill, New York.

Paškevičius, J. 1997. The Geology of the Baltic Republics. Vilnius University, Geological Survey of Lithuania, Vilnius.

Rõõmusoks, A. 1970. Стратиграфия вируской и харьюской серий (ордовик) Северной Эстонии, I (Stratigraphy of the Viruan Series (Middle Ordovician) in Northern Estonia, I). Valgus, Tallinn.

Rõõmusoks, A. 2004. Ordovician Strophomenoid Brachiopods of Northern Estonia. Fossilia Baltica, 3. University of Tartu, Tartu.

Ropot, V. F. and Pushkin, V. I. 1987. Ордовик Белоруссии (Ordovician of Belorussia). Nauka i tehnika, Minsk.

Schuchert, C. 1913. Class 2. Brachiopoda. In Textbook of Palaeontology (Zittel, K. A. von, ed.). 2nd ed. MacMillian, London.

Schuchert, C. and Cooper, G. A. 1932. Brachiopod Genera of the Suborders Orthoidea and Pentameroidea. Memoirs of the Peabody Museum of Natural History, 4(1). New Haven, Connecticut.
https://doi.org/10.5962/bhl.title.5732

Schuchert, C. and Cooper, G. A. 1956. Chazyan and Related Brachiopods, II. Smithsonian Miscellaneous Collections, 127(1).

Tsegelnyuk, P. D. 1976. Брахиоподы и стратиграфия нижнего палеозоя Волыно-Подолии (Brachiopods and Stratigraphy of Lower Palaeozoic of Volyn-Podollia). Naukova dumka, Kyiv.

Waagen, W. 1884. Salt Range fossils, vol. I, part 4. Productus Lime­stone fossils, Brachiopoda. Memoirs of the Geological Survey of India, Paleontologia Indica, 13(3–4), 547–728.

Williams, A. 1962. The Barr and Lower Ardmillan Series (Caradoc) of the Girvan District, South-West Ayrshire, with Description of the Brachiopoda.Geological Society of London Memoirs, 3, London.

Williams, A., Carlson, S. J., Brunton, C. H. C., Holmer, L. E. and Popov, L. E. 1996. A supra-ordinal classification of the Brachiopoda. Philosophical Transactions of the Royal Society of London (series B), 351(1344), 1171–1193.
https://doi.org/10.1098/rstb.1996.0101

Winchell, N. H. 1895. The Lower Silurian Brachiopoda of Minnesota. Final Report of Minnesota Geological Natural History Survey, 3, 333–474.

Back to Issue