Full understanding of diversity dynamics during the Great Ordovician Biodiversification Event (GOBE) requires analyses that investigate regional and species-level data and patterns. In this study, we combine bedding-plane scale data on brachiopod species counts and shell size collected from the Simpson Group of Oklahoma, USA, with species-level phylogenetic biogeography for three articulated brachiopod lineages that occurred throughout Laurentia. From these data, we ascertain that the primary influences of brachiopod shell size and diversity in the Simpson Group reflect global drivers, notably temporal position and paleotemperature. Similarly, the primary speciation pattern observed within Hesperorthis, Mimella, and Oepikina is the oscillation in speciation mode between dispersal and vicariance, which reflect the connection and disconnection of geographic areas, respectively. Processes that facilitate cyclical connectivity are global to regional in scale such as oceanographic changes, glacial cycles, or tectonic pulses. Therefore, both regional and continental scale analyses reinforce the importance of global factors in driving diversification during the GOBE.
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